By default, the DNS Resolver queries the root DNS servers directly and does not use DNS servers configured under System > General Setup or those obtained automatically from a dynamic WAN. This behavior may be changed, however, using the DNS Query Forwarding option. By contacting the roots directly by default, it eliminates many issues typically encountered by users with incorrect local DNS configurations, and the DNS results are more trustworthy and verifiable with Domain Name System Security Extensions (DNSSEC).
Further I am pretty curious about the dns behavior If I start using Unbound DNS: DNS over TLS - Does this overrule all other dns rel. settings and if in which way ?